Handbook of Agricultural Entomology

Handbook of Agricultural Entomology by Helmut van Emden is a landmark publication for students and practitioners of entomology applied to agriculture and horticulture. It can be used as a reference and as a general textbook. The book opens with a general introduction to entomology and includes coverage of the major insects (and mites) that cause harm to crops, livestock and humans. The important beneficial species are also included. Organisms are described in a classification of insect Orders and Families. The emphasis is on morphological characters of major taxonomic divisions, “spot characters” for the recognition of Families, and the life histories, damage symptoms and economic importance of the various pest species. The book is beautifully illustrated in full colour with more than 400 figures showing both the organisms and the damage caused to plants with diagnostic characters indicated by arrows. Coverage is world–wide and includes much material stemming from the vast personal experience of the author.

Companion Website details xv Preface xvii Acknowledgements xxi 1 The world of insects 1 1.1 The diversity of insects 1 1.2 The impact of insects on us 1 1.3 The impact we have on insects 5 1.4 Exploitation of insects 6 1.5 Other uses humans make of insects 6 1.6 Insect classification 7 2 External features of insects – structure and function 9 2.1 Introduction 9 2.2 The exoskeleton 10 2.3 The basic body plan of the insect 11 2.4 The head 11 2.5 The thorax 16 2.6 The abdomen 22 3 The major divisions of the Insecta 25 3.1 Introduction 25 3.2 Class Insecta, Subclass Apterygota or Phylum Arthropoda, Class Entognatha 25 3.3 Subclass Pterygota 26 4 Subclass Apterygota 31 4.1 Introduction 31 4.2 Order Diplura (two–pronged bristle–tails) 31 4.3 Order Protura 31 4.4 Order Thysanura (silverfi sh) 32 4.5 Order Collembola (springtails) 33 5 Subclass Pterygota, Division Exopterygota, Palaeopteran Orders 36 5.1 Introduction 36 5.2 Order Ephemeroptera (mayfl ies) 36 5.3 Order Odonata (dragonfl ies) 38 6 Subclass Pterygota, Division Exopterygota, Orthopteroid Orders 42 6.1 Introduction 42 6.2 Order Plecoptera (stoneflies) 43 6.3 Order Grylloblattodea 43 6.4 Order Mantophasmatodea (gladiators or heelwalkers) 44 6.5 Order Zoraptera (angel insects) 44 6.6 Order Orthoptera (grasshoppers and crickets) 44 6.7 Order Phasmida (stick and leaf insects) 51 6.8 Order Dermaptera (earwigs) 52 6.9 Order Embioptera (web spinners) 53 6.10 Order Dictyoptera (cockroaches and mantids) 54 6.11 Order Isoptera (termites) 57 7 Subclass Pterygota, Division Exopterygota, Hemipteroid Orders 63 7.1 Introduction 63 7.2 Order Psocoptera (booklice) 63 7.3 Order Mallophaga (biting lice) 64 7.4 Order Anoplura (= Siphunculata) (sucking lice) 64 7.5 Order Hemiptera (true bugs) 65 7.6 Order Thysanoptera (thrips or thunderflies) 106 8 Subclass Pterygota, Division Endopterygota, Lesser Orders 112 8.1 Introduction 112 8.2 Order Mecoptera (scorpion flies) 112 8.3 Order Siphonaptera (fleas) 113 8.4 Order Neuroptera 114 8.5 Order Trichoptera (caddis flies) 119 8.6 Order Strepsiptera (stylops) 120 9 Subclass Pterygota, Division Endopterygota, Order Lepidoptera (butterflies and moths) 124 9.1 Introduction 124 9.2 Suborder Glossata 128 10 Subclass Pterygota, Division Endopterygota, Order Diptera (true flies) 159 10.1 Introduction 159 10.3 Suborder Brachycera 173 10.4 Suborder Cyclorrhapha 174 11 Subclass Pterygota, Division Endopterygota, Order Hymenoptera (sawflies, ants, bees and wasps) 193 11.1 Introduction 193 11.2 Suborder Symphyta (sawflies) 194 11.3 Suborder Apocrita (ants, bees, wasps and parasitic wasps) 198 12 Subclass Pterygota, Division Endopterygota, Order Coleoptera (beetles) 221 12.1 Introduction 221 12.2 Suborder Adephaga 225 12.3 Suborder Polyphaga 231 13 Class Arachnida 272 13.1 Introduction 272 13.2 Subclass Araneida (spiders) 272 13.3 Subclass Acarina (mites and ticks) 274 Bibliography 284 Index 287

H. F. VAN EMDEN is an internationally respected entomologist who in the UK has been President of both the Royal Entomological Society and the Association of Applied Biologists. He is currently Emeritus Professor of Horticulture in the School of Agriculture, Policy and Development at the University of Reading, UK, where for 35 years he has taught entomology at Masters level to international students of Crop Protection. He has taught and carried out research on applied entomology in six continents.

“In summary, an excellent, well–illustrated handbook and a highly welcome addition to the literature of a very practical nature. ”  ( Experimental Agriculture , 1 January 2014) “Although the book might be too basic for researchers and professionals in the area, it is definitely a valuable resource for students and the general public. Summing Up: Highly recommended.  All undergraduate students and general readers.”  ( Choice , 1 October 2013) “The book is completed with a bibliography and index.  Overall it provides a very useful introduction to the range of insects that can be found in farming areas.”  ( International Pest Control , 1 July 2013) “For those with an interest in the birds of Iowa, raptors in general, or bird art.”  ( The Guardian , 1 April 2013) “Handbook of Agricultural Entomology by Helmut van Emden is a landmark publication for students and practitioners of entomology applied to agriculture and horticulture. It can be used as a reference and as a general textbook.”  ( Birdbooker Report , 15 April 2013) “Written by one of the world′s leading entomologists, Handbook of Agricultural Entomology is a landmark publication for students and practitioners of entomology applied to agriculture and horticulture. It can be used as a reference and as a general textbook.”  ( Phys.org , 18 March 2013)